240.15(B)(a) applies if the breaker opens via manual or manual operation (meaning, it opens all the hot wires whether it trips due to overload or someone shuts it off).
210.4(b) applies only to manually opening a breaker (i.e., to work on the circuit. If a handle-tie is used on two or more breakers for a MWBC, and one circuit trips, the others will stay on.