Again, not my area, but a couple of things came out from that (and they may be obvious to others)--
If they're properly hermetically sealed, no outside gasses should be able to get into the transformer. Does the gas pressure go negative when it cools after venting? Or... the gas has to be coming from somewhere, so if it's not being allowed into the transformer, it must already be there and is working its way out.
Oil is considered non-compressible, so gas pressure itself could not force the oil level down. It could force sidewalls out, lowering the level, but that ought to be unlikely. OTOH, if there are gas pockets somewhere in the transformer that should contain oil, that gas could compress and then the overall level would drop (like the classic u-tube manometer). With the transformer out of service, it might be interesting to pull a vacuum on the tank and see where it stabilizes and how the oil level changes.
(Were the transformers stored full? Was the oil changed before going into service? Did both transformers get about the same amount? Also gets back to the IR scan thing.)